Plants medium sized to large and robust, forming loose to dense mats or tufts. Stems spreading to ascending or erect, rarely subpendent, regularly to irregularly pinnately branched; in cross-section central strand present or absent; paraphyllia and pseudoparaphyllia present or absent. Leaves erect to wide spreading, falcate or not, oblong-lanceolate or broadly ovate to nearly oval, concave, plicate, smooth or somewhat plicate, apex broadly acute to rounded or gradually to abruptly acuminate, base cordate-auriculate or not; margins plane or recurved, entire to serrate or serrulate; costae double, 1/4-1/3(-1/2) lamina length or single; laminal cells smooth or some cells papillose at cell angles, medium cells oblong-linear, fusiform or linear, weakly porose or not; lower and basal cells elongate, strongly porose; alar cells differentiated or not. Dioicous. Perichaetia lateral. Setae elongate, smooth. Capsules erect to inclined or horizontal, urn ovoid-cylindrical to ellipsoid, ± curved and asymmetric; opercula conic to conic-apiculate or rostrate and oblique; peristome double, exostome teeth 16, smooth or cross-striate below, distally papillose, trabeculate on back; endostome lightly papillose, basal membrane high, segments 16, keeled and perforate, cilia 1-3, or absent to rudimentary. Calyptrae cucullate, naked and smooth. Spores spherical, finely papillose.