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Published In: Flora Taurico-Caucasica 3: 6. 1819. (Fl. Taur.-Caucas.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
General/Distribution: About 6 mainly psammophytic species in SE Europe, SW Asia to China. 1 recorded from Pakistan.


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Rigid, spiny dendroid-stellate pubescent annuals, with alternate flat, spine-tipped leaves. Flowers in axillary-cymes, bracteate, forming capitate or oblong-obovoid spiny glomerules borne in the axils of nearly all leaves. Perianth of (0-) 1-3 (-5) scarious segments. Stamens 1-5. Ovary with 2 stigmas. Fruit strongly flattened, opening at maturity at the front and back by means of scarious winglets, terminating in a long 2-cleft or 3-pronged beak. Seeds vertical, oval, compressed with semi-transparent endosperm; embryo circular.
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