Annual or perennial herbs, prostrate or erect to floating or scrambling, with entire, opposite leaves. Inflorescences of sessile or pedunculate heads or short spikes, axillary, solitary or clustered, bracteate. Flowers hermaphrodite, solitary in the axil of a bract, bibracteolate, bracts persistent but the perianth falling with the fruit, bracteoles persistent or not. Perianth segments 5, free, equal or unequal, glabrous or furnished with smooth or denticulate hairs. Stamens 2-5, some occasionally without anthers, the filaments distinctly monadelphous at the base into a cup or tube, alternating with large and dentate or laciniate to very small pseudo-staminodes (these rarely obsolete). Style short, stigma capitate. Ovary with a single pendulous ovule. Fruit an indehiscent utricle, thin-walled or sometimes corky; seeds ± lenticular.