Plants small to large, erect, dark‑green, green, or yellowish green in tufts or mats. Stems simple, forked or branching at base; stem epidermis of Mnium-type (Koponen 1968); rhizoids sparse and restricted to stem base or dense and occurring throughout the stem, macronemata rhizoids present, micronemata rhizoids absent. Leaves spirally arranged or distichous, linear‑lanceolate, elliptic or ovate; margins uni‑ or multistratose, bordered or elimbate, uni‑ or bi-serrate; costae single, strong, internally differentiated, smooth or toothed at back; upper cells short, hexagonal to isodiametric, alar cells not to weakly differentiated. Synoicous, autoicous, or dioicous. Perichaetia terminal, lateral, or basal. Setae elongate. Opercula rostrate. Peristome double. Calyptrae cucullate.