Plants mostly medium sized to rather large, forming loose mats or pendulous strands, rarely subdendroid, glossy green, yellowish- or reddish-brown. Stems pendent to spreading or suberect, irregularly to regularly pinnately branched; flagellate branches infrequent; in cross-section outer 3-6 rows of cells small, thick-walled, inner cells thin- to somewhat thick-walled, central strand present or more often absent; pseudoparaphyllia absent; rhizoids often clustered beneath primary stems. Leaves differentiated or not between stem and branch in size or shape, erect to erect-spreading, broadly ovate, concave, apex short acute to acuminate, base straight, rounded or auriculate; costae single and weak (occasionally forked) or absent; median cells linear, smooth, often porose; basal cells porose; insertion cells often golden-yellow; alar region weakly to strongly differentiated. Dioicous. Perichaetia lateral; leaves often differentiated. Setae somewhat elongate, smooth. Capsules exserted, erect to inclined, urn ovoid to broadly cylindrical; stomata usually present, at urn base, superficial; opercula rostrate; peristome double, exostome teeth 16, papillose; endostome basal membrane low to somewhat high, segments 16, often keeled and perforate, lightly papillose, cilia absent or rudimentary. Calyptrae mitrate, base not lobed, smooth, naked or hairy. Spores usually spherical, papillose.