Plants green, yellowish green to brown or dark brown, slender to robust, in loose or dense tufts. Primary stems creeping; secondary stems erect or prostrate, irregularly branched or pinnately branched; pseudoparaphyllia sometimes present. Leaves ovate at the base, concave, not plicate, gradually narrowed, with short or long slender apices, slightly decurrent at the base; margins plane, serrulate near the apex; costae single, ending below the apex; leaf cells oval or elliptical, regularly arranged; basal and marginal cells quadrate, thick-walled, smooth. Autoicous. Perigonia lateral; perichaetia on the tips of short branches; perichaetial leaves erect, inner perichaetial leaves larger, sheathing at the base, with or without costae; cells linear. Setae short; capsules erect, symmetric; opercula conic at the base, shortly apiculate, rarely shortly rostrate; annuli differentiated; stomata superficial, few; peristome mostly double; exostome teeth 16, lanceolate, pale yellowish, papillose, not striolate at the base, not distinctly trabeculate on inner surface; endostome segments linear-lanceolate or linear, rarely keeled or perforate; basal membrane low; cilia none. Calyptrae small, mitrate, rarely cucullate, somewhat roughened, rarely smooth. Spores spherical, medium-sized to large, smooth or papillose.