Plants small to large, whitish to pale green, grayish or pale brown plants in dense cushions; stems with central strand absent or weakly developed. Leaves thick and fleshy, consisting mostly of an expanded costa; in cross section with 2–several layers of large empty, hyaline, porose cells (leucocysts) enclosing a central layer of small, green cells (chlorocysts); laminae narrow, consisting of delicate, hyaline, quadrate, oblong or linear cells. Asexual reproductive structures sometimes present. Dioicous, pseudautoicous or autoicous. Setae terminal, straight; capsules erect or inclined, at times strumose; peristome absent or single and consisting of 8 or 16 lanceolate teeth. Calyptrae cucullate or mitrate, entire or fringed at base with long, single-celled cilia.