Plants erect or decumbent, small to robust. Stems simple or irregularly branched, often densely tomentose, hyalodermis present or absent, central strand present or absent. Leaves imbricate to wide-spreading or recurved, sheathing to erect-appressed at base or spreading from the insertion, smooth or plicate, straight, crisped, secund or circinate when dry, linear to variously lanceolate, leaf base strongly or not differentiated; costa single, strong, excurrent, often scabrous at back; upper leaf cells long-rectangular to quadrate, papillose, mammillose, prorulose, or smooth, alar cells not or well-differentiated. Synoicous, autoicous, or dioicous. Perigonial and perichaetial leaves sometimes long-linear. Setae short or long, straight or flexuose. Capsules cleistocarpous or stegocarpous, immersed to exserted, globose to oblong-cylindrical, erect to inclined, furrowed, wrinkled, rugose, or smooth; opercula nearly flat, plano-convex, conic-mammillate, or conic-rostrate; annuli weakly differentiated; peristome double, single or absent, exostome teeth papillose or smooth, dorsal trabeculae lightly thickened, ventral trabeculae strongly thickened, endostome rudimentary or well-developed, basal membrane high or low, segments broad, each segment split along the median line and each half diverging toward the cilia, cilia rudimentary to well-developed, sometimes fused. Spores spherical, oblong to reniform, often coarsely papillose. Calyptrae cucullate or conic.