Home Andean Bryophytes
Intro
Region
Name Search
Family List
Generic List
Andean Mosses XL database
Brymela Crosby & B.H. Allen Search in The Plant ListSearch in Index Nominum Genericorum (ING)Search in NYBG Virtual HerbariumSearch in JSTOR Plant ScienceSearch in SEINetSearch in African Plants Database at Geneva Botanical Garden Decrease font Increase font Restore font
 

Published In: Monographs in systematic botany from the Missouri Botanical Garden 11: 211. 1985. (Monogr. Syst. Bot. Missouri Bot. Gard.) Name publication detail
 

Project Name Data (Last Modified On 2/7/2013)
Acceptance : Accepted
Notes     (Last Modified On 2/7/2013)
Notes:
Brymela is distinguished by the rather large habit, the absence of or little developed stem hyalodermis, leaves lacking a border, with leaf margins serrated, of mostly simple teeth, and the ± homogeneous, firm- to thick-walled, porose laminal cells. A neotropical genus of some 12-13 species; in the tropical Andes found in wet lowland to premontane forest, rare at higher elevations; on logs, humus, leaf litter, and rocks. Our species were previously placed in Hookeriopsis.

 

Export To PDF Export To Word
Plants small to large and robust, forming loose to dense mats, glossy light or dark green to golden-brown. Stems and branches spreading to subascending; in cross-section central strand absent, hyalodermis absent. Leaves loosely erect-spreading to falcate-secund, oblong to oblong-short lanceolate, 0.8-8.0 mm long, smooth to undulate or rugose, apex acute, short acuminate or subobtuse, base weakly auriculate or not; margins reflexed or recurved in lower 1/3, or at midleaf, subentire to serrulate from about midleaf, often serrate at apex, elimbate or weakly limbate; costae double, 1/2-3/4 leaf length, parallel or slightly diverging, smooth to weakly or strongly serrulate along back; laminal cells smooth, apical cells shorter or not; median cells oblong-linear to linear, smooth to weakly or strongly porose; basal cells short, often golden-brown; marginal cells similar or forming a weak border of long linear cells. Asexual structures, absent. Perichaetia lateral, leaves differentiated, ovate-narrowly lanceolate, sharply serrate. Setae elongate, 10-45 mm long, slender to somewhat stout, smooth to more commonly weakly papillose or roughened below urn. Capsules inclined to horizontal, rarely subpendulous, urn ovoid-short cylindrical, 1-3 mm long, often somewhat flared below mouth when deoperculate; opercula, conic short to long rostrate; peristome, double, exostome teeth strongly furrowed, densely cross-striate, distal tips papillose; endostome basal membrane high, segments keeled and papillose. Calyptrae mitrate-short campanulate, naked, lobed at base. Spores spherical, lightly papillose.
 
Plantas pequeñas a grandes y robustas, formando tapices ralos a densos, verde claras brillantes o verde oscuras hasta pardo-doradas. Tallos y ramas extendidos a subascendentes; en sección transversal cordón central ausente, hialodermis ausente. Hojas laxamente erecto-extendidas a falcado-secundas, oblongas a oblongo-corto lanceoladas, 0.8-8.0 mm de largo, lisa a ondulada o rugosa, ápice agudo, corto acuminado o subobtuso, base levemente auriculada o no; márgenes reflexos o recurvados en 1/3 inferior, o en la mitad de la hoja, subenteros a serrulados desde cerca de la mitad de la hoja, generalmente aserrados en el ápice, elimbados o levemente limbados; costa doble, 1/2-3/4 del largo de la hoja, paralelas o ligeramente divergentes, lisas a levemente a fuertemente serruladas a lo largo del dorso; células de la lámina lisas, células apicales más cortas o no; células mediales oblongo-lineares a lineares, lisas a levemente o fuertemente porosas; células basales cortas, generalmente pardo-doradas; células marginales similares o formando un borde débil de células lineares largas. Estructuras asexuales, ausentes. Periquecios laterales, hojas diferenciadas, ovado-angostamente lanceoladas, fuertemente aserradas. Setas alargadas, 10-45 mm de largo, delgadas a algo gruesas, lisas a más comúnmente levemente papilosas o rugosas debajo de la urna. Cápsulas inclinadas a horizontales, raramente subpéndulas, urna ovoide-corto cilíndrica, 1-3 mm de largo, generalmente algo acampanada debajo de la boca luego de desoperculada; opérculo, cónico corto a largo rostrado; peristoma doble, dientes del exostoma fuertemente surcados, densamente estriados transversalmente, extremos distales papilosos; membrana basal del endostoma alta, segmentos plegados y papilosos. Caliptra mitrada-corta campanulada, desnuda, lobada en la base. Esporas esféricas, ligeramente papilosas.
 

Export To PDF Export To Word
1. Leaf apices acute to obtuse-rounded … 2.
1. Leaf apices acuminate …4.
2. Leaves falcate to falcate-secund, mostly less than 2 mm long… B. obtusifolia
2. Leaves loosely erect to erect-spreading, ± straight, not falcate, leaves mostly greater than 2 mm long … 3.
3. Leaf margins sharply serrate, teeth strongly bifid and swollen … B. rugulosa
3. Leaf margins weakly and bluntly serrulate, few teeth bifid but not swollen … B. parkeriana
4. Leaf apices cuspidate, with a notched sinuse at junction with lamina and leaf tip … B. cuspidata (and related forms)
4. Leaf apices gradually to± abruptly acuminate, neither cuspidate nor notched … 5.
5. Leaves 3.5-4 mm long … B. “antioquiana”
5. Leaves 2.4 mm long or less … 6.
6. Leaf apices short acuminate … B. acuminata
6. Leaf apices long and slenderly acuminate …  B. websteri
 
1. Apices foliares agudos a obtuso-redondeados … 2.
1. Apices foliares acuminados … 4.
2. Hojas falcadas a falcado-secundas, mayormente menores de 2 mm de largo … B. obtusifolia
2. Hojas laxamente erectas a erecto-extendidas, ± rectas, no falcadas, hojas mayormente mayores de 2 mm de largo … 3.
3. Márgenes foliares agudamente aserrados, dientes fuertemente bífidos e hinchados … B. rugulosa
3. Márgenes foliares levemente y obtusamente serrulados, con pocos dientes bífidos, pero no inflados … B. parkeriana
4. Apices foliares cuspidados, con un seno hendido en la unión entre la lámina y el ápice foliar … B. cuspidata (y formas relacionadas)
4. Apices foliares gradualmente a ± abruptamente acuminados, no cuspidados ni hendidos … 5.
5. Hojas 3.5-4 mm de largo … B. “antioquiana”
5. Hojas 2.4 mm de largo o menos … 6.
6. Apices foliares corto acuminados … B. acuminata
6. Apices foliares largos y delgadamente acuminados …  B. websteri
 
 
© 2014 Missouri Botanical Garden - 4344 Shaw Boulevard - Saint Louis, Missouri 63110