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Published In: Mémoires de la Société des Sciences Naturelles et Mathématiques de Cherbourg 16: 222. 1872. (Mém. Soc. Sci. Nat. Math. Cherbourg) Name publication detail

Project Name Data (Last Modified On 10/5/2009)
Acceptance : Accepted
Project Data     (Last Modified On 10/8/2017)
Plant Category: Mosses
General Reference:

Notes     (Last Modified On 10/8/2017)

      The genus is distinguished by the large, glossy plants; imbricate, concave, ecostate, acute to cuspidate ovate leaves; strongly auriculate base; weakly to strongly porose laminal cells; and excavate alar region. The following treatment is modified from Allen and Magill (2003).



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Plants light green to golden brown, mostly large and robust, forming loose mats or pendulous. Primary stems short to long creeping; rhizoids clustered beneath, rusty-red; leaves erect to appressed, broadly ovate-oval, ca. 2 mm long, to 1.6 mm wide, obtuse-rounded. Secondary stems often pendent or spreading, to 30 cm or more long; in cross-section 4–5 outer cell rows small, thick-walled and reddish-orange, inner cells large, ± thick-walled, pale yellow. Leaves imbricate, erect-spreading, stem and branch leaves similar, those of the stem often somewhat smaller, broadly ovate to oblong-ovate, to 3 mm long, deeply concave, acute and often cuspidate with tips mostly reflexed, base strongly auriculate and clasping; margins incurved, usually strongly so distally, entire or serrulate distally; costa none; laminal cells strongly to weakly porose or occasionally walls entire, median cells linear; alar region strongly differentiated, excavate, cells subquadrate to short rectangular, thick-walled, weakly porose, usually dark red or reddish-orange; basal cells golden-yellow. Dioicous. Perichaetial leaves to ca. 3.5 mm long, sheathing. Setae elongate, to 15 mm long, distally roughened, rusty-red. Capsules exserted, erect, urn ovoid to ovoid-short cylindrical (broadly ovoid when deoperculate), to 2 mm long; opercula long rostrate; peristome double (not observed). Calyptrae cucullate, hairy below, hairs ± erect. Spores not observed.



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Keys adapted from Allen & Magill (2003).


1. Plants with slender branchlets having microphyllous leaves, flagelliform branch tips, or deciduous leaves ... 2.

1. Plants lacking slender branchlets, flagelliform branches absent, leaves firm and nondeciduous ... 3.

2. Near all leaf apices erect, generally cuspidate to long cuspidate; flagelliform branch tips and/or deciduous leaves present ... P. mauiensis

2. Most leaf apices reflexed, mucronate; only deciduous leaves present ... P. flexilis form nudiramulosa

3. Near all leaf apices erect, generally cuspidate to long cuspidate ... P. mauiensis

3. Most leaf apices reflexed, obtuse, broadly acute, mucronate or apiculate ... 4

4. Plants medium-sized to robust, leaves 1.7–3.0 mm long ... P. flexilis

4. Plants small to medium-sized, leaves 1.2–1.6 mm long ... P. vermiformis



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