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Published In: Botanische Zeitung (Berlin) 28(3): 35. 1870. (21 Jan 1870) (Bot. Zeitung (Berlin)) Name publication detailView in Biodiversity Heritage Library

Project Name Data (Last Modified On 9/18/2009)
Acceptance : Accepted
Project Data     (Last Modified On 10/8/2017)
Plant Category: Mosses

Notes     (Last Modified On 10/8/2017)

Mittenothamnium, as presently recognized, is similar to Chryso-hypnum but differs in having a primary stipe-like stem, lanceolate pseudoparaphyllia, and leaf cells usually projecting only at distal angles. Chryso-hypnum segregated from Mittenothamnium, is poorly known. In both genera there are an inflated number of species described mostly from the Neotropics, and less so from Africa and associated islands. There may be only 5-10 species. The genus was previously placed in Mittenothamnium, and earlier in Microthamnium and Stereohypnum. Keys adapted from Nishimura and Ando (in Sharp et al. 1994).


Mittenothamnium es similar a Chryso-hypnum pero se diferencia en tener un tallo primario como estípite, pseudoparáfilos lanceolados, y células de las hojas generalmente proyectándose solo en los ángulos distales. Chryso-hypnum separado de Mittenothamnium, es un género poco conocido. En ambos géneros hay un número inflado de especies descritas generalmente del Neotrópico y menos de áfrica e islas asociadas. Debe haber tan solo 5-10 especies.


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Plants small, forming mats, pale to rather dark green, or yellow. Stems procumbent, distally stems and branches spreading to ascending, subpinnately to pinnately branched, branches short; in cross-section with outer 2-3 rows of cells small, thick-walled, golden-yellow to rusty-red, inner cells larger, thin-walled, central strand absent; pseudoparaphyllia filamentous to subfilametous; rhizoids clustered beneath, rusty red. Stem leaves ovate-lanceolate, 0.6-0.9 mm long, to 0.35 mm wide, apex acuminate, often narrowly so, straight to curved; margins plane above, reflexed near base, serrulate in distal 2/3 or more, dentate below; costae short and forked or appearing single, ca. 1/5 lamina length, occasionally absent; median cells oblong-linear, both lower and upper angles projecting; alar region somewhat strongly to weakly differentiated, cells few, quadrate to short rectangular. Branch leaves similar, often complanate, progressively smaller. Autoicous. Perigonia to 0.4 mm long. Perichaetial leaves ovate-subulate, to 1.2 mm long. Seta 9-30 mm long, slender. Capsule pendent, urn ovoid, ca.0.6-1.0 mm long, curved; exothecial cells irregularly short rectangular to subquadrate, thick-walled; operculum conic-apiculate; peristome with exostome teeth finely striate below, distally papillose; endostome basal membrane high, segments keeled, perforate, papillose, cilia 2-3. Calyptra naked or with few hairs. Spores lightly papillose.


Plantas pequeñas, formando matas, verdes pálidas hasta bastante oscuras, o amarillas. Tallos procumbentes, tallos y ramas distales patentes hasta ascendentes, ramificaciones subpinnadas hasta pinnadas, ramas cortas; en corte transversal con 2-3 filas externas de células pequeñas, con paredes gruesas, amarillo-doradas hasta rojas-oxidadas, células internas mayores, con paredes delgadas, cordón central ausente; pseudoparáfilos filamentosos hasta subfilamentosos; rizoides agregados abajo, rojos ferrugíneos. Hojas del tallo ovado-lanceoladas, 0.6-0.9 mm de largo, hasta 0.35 mm de ancho, ápices acuminados, a menudo angostamente, rectas hasta curvadas; márgenes planos arriba, reflexos cerca de la base, serrulados hasta 2/3 o más arriba, dentados abajo; costa corta y bifurcada o de apariencia simple, ca. 1/5 de la longitud de la lámina, ocasionalmente ausente; células medias oblongo-lineales, con proyecciones en los ángulos inferiores y superiores; región alar un poco marcada hasta débilmente diferenciada, pocas células, cuadradas hasta corto-rectangulares. Hojas de las ramas similares, a menudo complanadas, progresivamente menores. Autoicas. Perigonio de 0.4 mm de largo. Hojas periqueciales ovado-subuladas, hasta 1.2 mm de largo. Setas 9-30 mm de largo, delgadas. Cápsulas pendientes, urna ovoide, ca. 0.6-1.0 mm de largo, curvada; células exoteciales irregularmente corto-rectangulares hasta subcuadradas, con paredes gruesas; opérculo cónico-apiculado; peristoma con dientes del exostoma finamente estriados abajo, distalmente papilosos; membrana basal del endostoma alta, segmentos carinados, perforados, papilosos, 2-3 cilios. Caliptra desnuda o con pocos pelos. Esporas finamente papilosas.


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1. Stem leaves straight-acuminate; alar cells quadrate to subquadrate in 3-6 rows; setae to 2 cm long … C. diminutivum
1. Stem leaves curved-acuminate; alar cells only weakly differentiated, short-rectangular; setae more than 2 cm long … C. elegantulum
1. Hojas de los tallos rectas-acuminadas; células alares cuadradas a subcuadradas en 3-6 filas; setas hasta 2 cm de largo … C. diminutivum
1. Hojas de los tallos curvadas-acuminadas; células alares sólo débilmente diferenciadas, corto-rectangulares; setas más de 2 cm de largo … C. elegantulum
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