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Published In: Bryologia Universa 1: 480. 1826. (Bryol. Univ.) Name publication detailView in Biodiversity Heritage Library

Project Name Data (Last Modified On 9/20/2009)
Acceptance : Accepted
Project Data     (Last Modified On 10/8/2017)
Plant Category: Mosses
General Reference:

Notes     (Last Modified On 10/8/2017)
Ceratodon contains 4-5 species distributed worldwide; two species in the Neotropics. The genus in the tropical Andes is typically found in open montane, páramo and puna, often in disturbed and cleared sites; on soil or rock, occasionally on logs or rarely on tree trunks. Burley and Pritchard (1990) have recognized C. stenocarpus Bruch & Schimp. as a subspecies of C. purpureus (Hedw.) Brid.; in the present treatment these two taxa are retained as separate species. In the Neotropics C. stenocarpus is by far the more common taxon. Many collections determined as C. purpureus are C. stenocarpus.
Ceratodon contiene 4-5 especies distribuidas en todo el mundo; dos especie en el Neotrópico. En los Andes tropicales este género se encuentra típicamente en bosque montano abierto, páramo y puna, a menudo en sitios alterados y desmontados; sobre suelo o rocas, ocasionalmente sobre troncos en descomposición o rara vez sobre troncos de árboles.


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Plants small to medium sized, forming loose to dense tufts, dark green to blackish-green. Stems erect, several branched by innovations; outer 1-2 rows of cells small, thick-walled, inner cells progressively larger, thin-walled, central strand present; radiculose below. Leaves lanceolate or triangular-lanceolate, keeled distally, apex short acuminate; margins reflexed, entire to distally dentate or irregularly serrate at apex; costae single, strong, subpercurrent to short excurrent, in cross-section stereids above and below guide cells; median cells quadrate to subquadrate, smooth, thick-walled; basal cells rectangular, thick-walled. Dioicous. Perichaetia terminal; leaves larger, sheathing at base. Setae elongate, smooth. Capsules suberect to horizontal, urn short to long ovoid-cylindrical, weakly to strongly curved, asymmetric, furrowed when deoperculate and dry; exothecial cells irregularly long rectangular, thin-walled; stomata few at urn base, superficial; annuli in 2-3 rows, revoluble; opercula conic; peristome single, teeth 16, divided 2/3 or more to near base, thickened segments jointed or not, papillose. Calyptrae cucullate, smooth and naked. Spores spherical, lightly papillose.
Plantas pequeñas a medianas, formando manojos laxos a densos, de color verde oscuro a verde-negruzco. Tallos erectos, muy ramificado por innovaciones; 1-2 filas externas de células pequeñas con paredes gruesas, células internas progresivamente mayores, con paredes delgadas, cordón central presente; radiculosos abajo. Hojas lanceoladas o triangulares-lanceoladas, distalmente carinadas, ápices corto-acuminados; márgenes reflexos, enteros hasta distalmente dentados o irregularmente serrados en el ápice; costa simple, fuerte, subpercurrente hasta corto-excurrente, en corte transversal estereidas encima y debajo de las células guía; células mediales cuadradas a subcuadradas, lisas, con paredes gruesas; células basales rectangulares con paredes gruesas. Dioicas. Periquecios terminales; hojas mayores, envainadoras en la base. Setas alargadas y lisas. Cápsulas suberectas hasta horizontales, urna corta hasta larga ovoide-cilíndrica, débilmente hasta marcadamente curvadas, asimétricas, acanaladas cuando están desoperculadas y secas; células exoteciales irregularmente largo-rectangulares, con paredes delgadas; pocos estomas en la base de la urna, superficiales; anillo en 2-3 filas, revolutos; opérculos cónicos; peristomas simples, con 16 dientes, 2/3 partidos o más cerca de la base, segmentos engrosados articulado o no, papilosos. Caliptra cuculada, lisa y desnuda. Esporas esféricas, finamente papilosas.

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1. Peristome teeth cross-bared in lower 1/2-1/3, appearing perforate, distal tips finely papillose; seta purple; capsule horizontal, ovoid, sulcate when old … C. purpureus
1. Peristome teeth separate to near base or cross-bared just above base, tips coarsely papillose or papillose-spiculose; seta pale-yellow or red; capsule suberect, cylindric … C. stenocarpus
1. Dientes del peristoma con bandas transversales 1/2-1/3 basales aparentemente perforados, extremos distales finamente papilosos; seta púrpura; cápsula horizontal, ovoide, sulcada en la madurez … C. purpureus
1. Dientes del peristoma separados hasta casi la base o con bandas transversales solo arriba de la base, extremos ásperamente papilosos o papiloso-espiculosos; seta amarillo-pálida o roja; cápsulas suberectas, cilíndricas … C. stenocarpus
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