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Published In: Muscologiae Recentiorum Supplementum 4: 71. 1819[1818]. (Muscol. Recent. Suppl.) Name publication detail

Project Name Data (Last Modified On 4/11/2011)
Acceptance : Accepted
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Campylopus is a large, taxonomically complex group of worldwide distribution. Due to great variation in nearly all its characters the generic limits of the genus are diffuse. In general, Campylopus consists of medium to large, dioicous mosses with stiff, erect to spreading (generally unbordered), linear leaves having broad, excurrent costa approximately 1/2 to 2/3 the leaf base width. Many species have costae with a ventral layer of variously enlarged hyaline cells (hyalocysts). Sporophytically, the cygneous setae, capsules that lack stomata, and dicranaeous peristome teeth are distinctive. The calyptrae are cucullate and frequently ciliate at base.

In the typical sense Campylopus is easy to recognize, but the genus harbors species whose characters grade into those of Atractylocarpus, Dicranodontium, Dicranella, Campylopodium, Paraleucobryum, and Pilopogon (see Allen 1989). In the following treatment Campylopus is divided into two groups on the basis of the form of the costal cells ventral to the guide cells.


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Campylopus Brid., Musc. Recent. Suppl. 4: 71. 1819.

Plants small to robust, densely tufted; stems sparsely branched, often densely tomentose; leaf rhizoids frequently on the lower dorsal surface of the costa; in cross section with a well-developed central stand. Leaves ovate-lanceolate to ovate-subulate, variably toothed, nearly always denticulate at the extreme apex; costa broad, occupying most of the upper leaf, percurrent to excurrent, often ribbed or mammillose at back, at times with multi-cellular lamellae; cells smooth, porose or entire, upper cells quadrate, oval, rhombic, oblong, rhomboidal or elongate, usually in oblique rows, incrassate, marginal cells at base at times forming a hyaline border; alar cells commonly well-developed. Dioicous. Setae cygneous when moist; capsules exserted, ovoid to ellipsoid, strumae present or absent, erect or curved; stomata absent; annuli present; peristome teeth 1/2 to nearly their full lengths divided into two filaments, generally striate on the outer surface; opercula rostrate. Calyptrae cucullate, entire or fringed at base by long, single-celled hairs. Spores yellow to brown, variously papillose.



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