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Discussion:
Acidodontium, with 11
species (Ochi 1992), is restricted to the neotropics. The gametophytes of Acidodontium
are nearly inseparable from some large species of Bryum or Brachymenium.
The genus is based on sporophytic features. Its capsules are generally erect
and have long, slender, abruptly constricted necks, the operculum is
hemispheric and only occasionally apiculate. Its exostome teeth are broad in
the lower _ then abruptly constricted to slender filaments, and the dorsal
(outer) trabeculae project laterally, i.e., in the direction of the
adjacent teeth. The endostome in Acidodontium has a high basal membrane
with broad segments that are completely split along the keel and laterally
divergent in the direction of the adjacent segments. Although the above suite
of sporophytic characters neatly circumscribes the genus, a related form of its
odd endostome is found in Brachymenium columbicum where the segments are
split along the keel and divergent, but in this case each ½ segment is fully
united with the adjacent cilia and ½ segment. An endostome nearly identical to
that of Acidodontium is also found in the Bartramiaceae (Bartramia,
Breutelia, Philonotis, and Plagiopus).
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